(1)

(2)

Fig. 1
(a) A graph whose vertices (squares) are connected by unidirectional edges (arrows). (b) Network structure corresponding to graph (a). The black solid and dashed arrows indicate excitatory links corresponding to systems (1) and (2), respectively. The gray solid and dashed lines indicate inhibitory links, which are added due to (C4). (c) (C1)(C2) Regulatory rules for excitatory and inhibitory interactions. (C3) Transient behavior when a node receives no input. The solution converges to an incoherent state for any initial condition. (C4) Bidirectional inhibitory interactions take place at the branching point. The activity of the inhibitory interactions is also controlled by s m (dashed line)








The systems (1) and (2) are decoupled by taking
. The former and latter systems have forward and opposite directional excitatory links. That is, if a link from
exists for (1) (
), then a link from
exists for (2) (
). System (1) with
is the same as the system proposed in [10], and system (2) with
is the same as the previous system, except for the direction of the links. We refer to system (1) with
and system (2) with
as system A and system B, respectively. We refer to the coupled system with
as system C.










The parameters p, q, r, and
are independent of k ±, and set p = 0. 02, q = 1. 0,
, and
. Furthermore, w(t) is a small amount of random noise in the interval [0, 0. 05]. The time constants τ j take random values from the interval between τ − and τ +, where the values of τ − and τ + are set to
. The distribution of τ j is the same for all nodes; that is, it is independent of k.





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